The Knights of Destruction attack the Fortress of Infinity, while Nerissa, posing as the Mage, corrupts Halinor Clarkson, the former Guardian of Fire. By accident, Cornelia receives the power of a Quinto Guardian, possessing all five elements, and Nerissa tries to use this to her advantage. W.I.T.C.H. must clean the cars and Will is reunited with her father.

While Irma is trying to organize a concert for hit singer Vance Michael Justin, Caleb learns that Nerissa is his mother and confronts her in her hideaway at Mount Thanos. The girls struggle with Irma, who is giving them many orders to help organize a concert.

Nerissa and her Knights of Destruction attack Zamballa, a world of purple foliage with a populace of moving trees, disguised as the Guardians. When W.I.T.C.H. arrives to stop them, they are forced to fend off the Zamballians and their leader, Queen Kadma Starlighter, the former Guardian of Earth. Taranee starts to be angry about her mother.