When a snowstorm blocks Kate from accessing the zoo, Melman decides to become "substitute Kate" and make sure the animals are cared for; a series of stumbles makes Melman question whether he's cut out for the job.

Gloria meets a shy hermit crab with a secret talent for tap dancing and plans a flashy production; when she pushes the introverted crab too far, she learns to respect her shy friends' comfort levels.

When Lala announces she's moving out of the habitat, Gloria feels sad. But instead of admitting it, she puts all her energy into throwing the "happiest" goodbye party for her tank mate's last day in the habitat.