After growing into a froglet, Lala is ready to finally join the gang on a big adventure in the city; Gloria goes overboard in trying to protect her tiny friend in the big world -- only to realize that she underestimated Lala.

When a snowstorm blocks Kate from accessing the zoo, Melman decides to become "substitute Kate" and make sure the animals are cared for; a series of stumbles makes Melman question whether he's cut out for the job.

Gloria meets a shy hermit crab with a secret talent for tap dancing and plans a flashy production; when she pushes the introverted crab too far, she learns to respect her shy friends' comfort levels.