The six friends gather for a wedding; Brewster suspects Will of subterfuge; problems arise with Carla's job and one promised to her by Jenna; and an old truth throws Detective Marjorino for a loop.

Craig struggles to keep his dad's company afloat; Will prepares for international travel; Carla gets a title Sam covets, and a warning; Jenna comes home to visit her ill mother; and Aaron finds inspiration in Prague.

In 1991, Will visits Jenna in Hollywood en route to completing a mission; Craig is disappointed by both Aaron and Sam; and in 2005, the gun resurfaces and another cryptic note is received, this time by Will.