Bill Richardson has his hands full representing the McCullough's in the custody case, much to Izzy's dismay. Lexie finds out upsetting news and turns to Pearl for help. Meanwhile, Elena – feeling as though this whole custody battle is her fault – goes to New York to search for clues about Mia's past and enlists the help of an old acquaintance.

In 1981, a young Mia begins studying at the School of Visual Arts in New York City, where she meets the captivating photographer Pauline Hawthorne. Struggling to pay her tuition, Mia makes a decision that will change the course of her entire life. Meanwhile, in Ohio, a young Elena questions past choices as she adjusts to life with a fourth child.

Mia, Bebe, and lawyer Ed Lan prepare their case against the McCulloughs. Izzy grows closer to Mia over a shared experience while her relationship with Elena reaches a boiling point. Bill and Elena's marriage is tested. Trip and Pearl continue to explore their relationship while Lexie and Brian consider ending theirs.