Having been transported into the future, Buffy joins forces with Melaka Fray, a fellow-slayer whose twin brother, Harth, is a vampire contemplating an evil plan. Meanwhile, back in the present, Xander and Dawn struggle to defend the castle.

While Willow seeks otherworldly help in bringing Buffy back from the future, Xander and Dawn flee deep into the woods, where they encounter a strange race of tree-like forest souls. Elsewhere, in the future, after taking on a horde of vampires, Fray strikes a deal with Dark Willow.

Xander and Dawn lead an army of wood creatures into battle against Warren and Amy's minions. And, with a potentially cataclysmic temporal rift about to open, Buffy must escape from Dark Willow and defeat Fray in order to return to the present and save the world.