Si King and Dave Myers explore Oxfordshire, where they resurrect a lost county recipe and cook it for the locals in Henley. They make cheese with rock star Alex James and sample the local quince. Finally, they face a cook-off against top Oxfordshire chef Emily Watkins. Restaurant diners decide whose dish best defines the taste of their county.

Si King and Dave Myers explore East Sussex, where they cook a hearty dish on Brighton beach. They taste local sparkling wine every bit as good as champagne and pay a visit to the Rye Scallop Festival. Finally, they face a cook-off against top chef Ross Pavey. Restaurant diners decide whose dish best defines the taste of East Sussex in a blind tasting.

Si King and Dave Myers explore Gwynedd, where they cook a traditional county favourite in Portmeirion. They visit a mountain lamb farmer and meet a man growing exotic mushrooms in shipping containers. Finally, they face a cook-off against top chef Peter Jackson. Restaurant diners decide whose dish best defines the taste of Gwynedd in a blind tasting.