Si King and Dave Myers explore County Antrim, where they cook a traditional county favourite at Carrickfergus Castle, pick potatoes and head to the coast in search of dulse seaweed. Finally, they use the best local ingredients in a cook-off against top chef Niall McKenna. Restaurant diners decide in a blind tasting who has created the dish that best defines their county.

Si King and Dave Myers explore Oxfordshire, where they resurrect a lost county recipe and cook it for the locals in Henley. They make cheese with rock star Alex James and sample the local quince. Finally, they face a cook-off against top Oxfordshire chef Emily Watkins. Restaurant diners decide whose dish best defines the taste of their county.

Si King and Dave Myers explore East Sussex, where they cook a hearty dish on Brighton beach. They taste local sparkling wine every bit as good as champagne and pay a visit to the Rye Scallop Festival. Finally, they face a cook-off against top chef Ross Pavey. Restaurant diners decide whose dish best defines the taste of East Sussex in a blind tasting.