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In 2016, a meltdown of a nuclear power plant contaminates Tokyo, forcing the government to order its citizens to evacuate. 20 years later, Tokyo has become a ghost town due to the high levels of radiation which the government has banned entry for anyone. When a distress signal is received from the area, the Japan Ground Self-Defense Force dispatches three girls from the special unit Coppelion to search the ruined capital for survivors.